Mammals (formally Mammalia) are a class of vertebrate animals whose females are characterized by the possession of mammary glands while both males and females are characterized by sweat glands, hair, three middle ear bones used in hearing, and a neocortex region in the brain.

Except for the five species of monotremes (which lay eggs), all mammal species give birth to live young. Most mammals also possess specialized teeth, and the largest group of mammals, the placentals, use a placenta during gestation. The mammalian brain regulates endothermic and circulatory systems, including a four-chambered heart.

There are approximately 5,400 species of mammals, distributed in about 1,200 genera, 153 families, and 29 orders^[1] (though this varies by classification scheme). Mammals range in size from the 30–40-millimetre (1.2–1.6 in) Bumblebee Bat to the 33-metre (110 ft) Blue Whale.

Mammals are divided into two subclasses, the prototheria, which includes the egg-laying monotremes, and the theria, which includes the live-bearing marsupials and placentals. Most mammals, including the six largest orders, belong to the placental group. The three largest orders, in descending order, are Rodentia (mice, rats, and other small, gnawing mammals), Chiroptera (bats), and Soricomorpha (shrews, moles and solenodons). The next three largest orders include the Carnivora (dogs, cats, weasels, bears, seals, and their relatives), the Cetartiodactyla (including the even-toed hoofed mammals and the whales) and the Primates to which the human species belongs. The relative size of these latter three orders differs according to the classification scheme and definitions used by various authors.

Phylogenetically, Mammalia is defined as all descendants of the most recent common ancestor of monotremes (e.g., echidnas and platypuses) and therian mammals (marsupials and placentals).^[2] This means that some extinct groups of "mammals" are not members of the crown group Mammalia, even though most of them have all the characteristics that traditionally would have classified them as mammals.^[3] These "mammals" are now usually placed in the unranked clade Mammaliaformes.

The mammalian line of descent diverged from an amniote line at the end of the Carboniferous period. One line of amniotes would lead to reptiles, while the other would lead to synapsids, including mammals. The first true mammals appeared in the Jurassic period. Modern mammalian orders appeared in the Palaeocene and Eocene epochs of the Palaeogene period.

Distinguishing features

Living mammal species can be identified by the presence of sweat glands, including those that are specialized to produce milk.

However, other features are required when classifying fossils, since soft tissue glands and some other features are not visible in fossils. Paleontologists use a distinguishing feature that is shared by all living mammals (including monotremes), but is not present in any of the early Triassic synapsids: mammals use two bones for hearing that were used for eating by their ancestors. The earliest synapsids had a jaw joint composed of the articular (a small bone at the back of the lower jaw) and the quadrate (a small bone at the back of the upper jaw). Most reptiles and non-mammalian synapsids use this system including lizards, crocodilians, dinosaurs (and their descendants the birds), and therapsids (mammal-like "reptiles"). Mammals have a different jaw joint, however, composed only of the dentary (the lower jaw bone which carries the teeth) and the squamosal (another small skull bone). In mammals the quadrate and articular bones have become the incus and malleus bones in the middle ear. Note: "non-mammalian synapsids" above implies that mammals are a sub-group of synapsids, and that is exactly what cladistics says they are.

Mammals also have a double occipital condyle: they have two knobs at the base of the skull which fit into the topmost neck vertebra, and other vertebrates have a single occipital condyle. Paleontologists use only the jaw joint and middle ear as criteria for identifying fossil mammals, since it would be confusing if they found a fossil that had one feature, but not the other.

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